Heracleum barbatum var. turugisanense (Honda) M.Hiroe variety synonym

Wendia argaea (Boiss. & Balansa) Woronow species synonym

Wendia schelkovnikowii (Woronow) Grossh. species synonym

Sphondylium lanatum var. angustum Nakai variety synonym

Heracleum lanatum subsp. akasimontanum (Koidz.) Kitam. subspecies synonym

Heracleum pubescens var. glabratum Sommier & Levier variety synonym

Heracleum pastinacifolium subsp. schelkovnikowii (Woronow) Satzyp. subspecies synonym

Heracleum moellendorffii var. turugisanense (Honda) Ohwi variety synonym

Wendia incana var. lazica (Boiss.) Grossh. variety synonym

Heracleum sphondylium f. dissectum H.Ohba form proparte synonym

Heracleum mantegazzianum × sphondylium subsp. sphondylium species accepted

Heracleum mantegazzianum x persicum species synonym

Heracleum persicum x sphondylium subsp. sibiricum species synonym

Heracleum persicum x sphondylium ssp. sibiricum species synonym

Heracleum mantegazzianum x Heracleum sphondylium species accepted

Heracleum mantegazzianum × persicum species accepted

Heracleum mantegazzianum x sphondylium ssp. sphondylium species synonym

Heracleum persicum × sphondylium ssp. sibiricum species synonym

Wendia schelkovnikowii (Woronow) Woronow species synonym

Heracleum montegazzianum Sommier & Levier species synonym

Heracleum leichtlinii J.N.Haage & E.Schmidt species doubtful

Heracleum atlanticum (Coss. ex Batt.) T.Durand & B.D.Jacks., 1902 species accepted

Heracleum sphondylium var. turugisanense (Honda) H.Ohba variety synonym

Heracleum ternatum var. angustisectum Stoj. & Acht. variety synonym

Heracleum angustisectum (Stoj. & Acht.) Peev species accepted

Heracleum calcareum subsp. colchicum (Lipsky) Satzyp. subspecies synonym

Pastinaca schelkovnikowii (Woronow) Koso-Pol. species synonym

Heracleum piliferum Kljuykov & Lyskov species accepted

Heracleum kurdistanicum Rastegar, Maroofi & Tabad species accepted

Heracleum sphondylium var. austriacum (L.) Fiori variety synonym

Pastinaca sphondylium var. sibiricum (L.) Fiori variety synonym

Sphondylium proteiforme Crantz species synonym

Heracleum alpinum f. heterophyllum Charpié form homotypic synonym

Heracleum sphondylium f. heterophyllum (Charpié ex Thell.) Hegi form synonym

Heracleum sphondylium f. puberulum Thell. form synonym

Sphondylium giganteum Fisch. species homotypic synonym

Heracleum persicum var. glabrescens (Boiss. & Hohen.) Thell. variety synonym

Heracleum speciosum var. pubescens (Hoffm.) Thell. variety synonym

Heracleum giganteum Fisch. ex Spreng. species synonym

Heracleum alpinum f. heterophyllum Charpié ex Thell. form synonym

Heracleum mantegazzianum f. rosicalyx Schwer. form synonym

Selinum persicum (Desf. ex Fisch., C.A.Mey. & Avé-Lall.) E.H.L.Krause species synonym

Heracleum schelkovnikowii Woronow species accepted

Heracleum schansianum Fedde species homotypic synonym

Heracleum scabrum Albov species accepted

Heracleum scabridum Franch. species accepted

Heracleum roseum Steven species accepted

Heracleum ×rodnense Nyar. & Todor species accepted

Heracleum rechingeri Manden. species accepted

Heracleum rawianum C.C.Towns. species accepted

Heracleum rapula Franch. species accepted

Heracleum pusillum Pall. ex Ledeb. species synonym

Heracleum pulchrum Zumagl. species doubtful

Heracleum pulchellum Pall. ex Ledeb. species synonym

Heracleum pubescens (Hoffm.) M.Bieb. species accepted

Heracleum ponticum (Lipsky) Schischk. ex Grossh. species accepted

Heracleum platytaenium Boiss. species accepted

Heracleum persicum Desf. species accepted

Heracleum pastinacifolium subsp. transcaucasicum (Manden.) P.H.Davis subspecies synonym

Heracleum pastinacifolium subsp. incanum (Boiss. & A.Huet) Davis subspecies synonym

Heracleum pastinaca Fenzl species accepted

Heracleum paphlagonicum Czeczott species accepted

Heracleum panaces Gueldenst. species heterotypic synonym

Heracleum palmatifidum Fisch. & Avé-Lall. species synonym

Heracleum pachyrhizum Sommier & Levier species synonym

Heracleum ossethicum Manden. species accepted

Heracleum orphanidis Boiss. species accepted

Heracleum oreocharis H.Wolff species accepted

Heracleum oncosepalum (Hand.-Mazz.) Pimenov & Kljuykov species accepted

Heracleum nipponicum Kitag. species synonym

Heracleum nanum Satzyp. species synonym

Heracleum morifolium H.Wolff species synonym

Heracleum montanum Georgi ex Ledeb. species heterotypic synonym

Heracleum moellendorffii var. sagenifolium K.T.Fu variety synonym

Heracleum moellendorffii var. paucivittatum R.H.Shan & T.S.Wang variety synonym

Heracleum moellendorffii var. akasimontanum (Koidz.) Okuyama ex T.Shimizu variety synonym

Heracleum minimum Lam. species synonym

Heracleum microcarpum Franch. species synonym

Heracleum massyciticum Stapf & Wettst. species synonym

Heracleum marginatum Fisch. ex Hoffm. species synonym

Heracleum mandenovae Satzyp. species synonym

Heracleum longifolium Hegetschw. species synonym

Heracleum likiangense H.Wolff species accepted

Heracleum ligusticifolium M.Bieb. species accepted

Heracleum leichtlinii Vatke species doubtful

Heracleum lehmannianum Bunge species accepted

Heracleum lanceolatum Bordz. species synonym

Heracleum lanatum f. rubriflorum (H.Ohba) T.Yamaz. form synonym

Heracleum lanatum f. dissectum (H.Ohba) T.Yamaz. form synonym

Heracleum lallemantii species accepted

Heracleum laciniatum Hornem. species synonym

Heracleum kansuense Diels species accepted

Heracleum juranum Genty species synonym

Heracleum jugatum Boiss. species doubtful

Heracleum jacquemontii C.B.Clarke species accepted

Heracleum isopetalum Maxim. species synonym

Heracleum insigne Dalla Torre & Sarnth. species doubtful

Heracleum inperpastum Koidz. species synonym

Heracleum inexpectatum C.Norman species synonym

Heracleum incanum Boiss. & A.Huet species accepted

Heracleum idae Kulieva species synonym

Heracleum humile Sm. species accepted

Heracleum henryi H.Wolff species accepted

Heracleum hemsleyanum Diels species accepted

Heracleum gummiferum Willd. species synonym

Heracleum grossheimii Manden. species synonym

Heracleum graveolens (C.A.Mey.) species accepted

Heracleum grandiflorum Steven ex M.Bieb. species accepted

Heracleum gorganicum Rech.fil. species accepted

Heracleum glabrescens Boiss. & Hohen. species synonym

Heracleum giganteum Fisch. ex Hornem. species synonym

Heracleum freynianum Sommier & Levier species accepted

Heracleum franchetii M.Hiroe species accepted

Heracleum forrestii H.Wolff species accepted

Heracleum fargesii H.Boissieu species accepted

Heracleum eminens Lange species synonym

Heracleum elgonense (H.Wolff) Bull. species accepted

Heracleum egrissicum Gagnidze species accepted

Heracleum dulce var. akasimontanum (Koidz.) Kitag. variety synonym

Peucedanum kirungae (Engl.) M.Hiroe species synonym

Heracleum douglasii DC. species synonym

Heracleum dissectum var. voroschilovii (Gorovoj) Pimenov variety synonym

Heracleum dissectum subsp. moellendorffii (Hance) Vorosch. subspecies synonym

Heracleum dissectifolium K.T.Fu species accepted

Heracleum dichotoma Benth. & Hook.f. species doubtful

Heracleum decipiens Fisch. ex Hoffm. species synonym

Heracleum davisii Manden. species synonym

Heracleum dalgadianum S.M.Almeida species accepted

Heracleum cyclocarpum C.Koch species doubtful

Heracleum cuneiforme DC. species doubtful

Heracleum cuneatilobatum Hausskn. & Bornm. species synonym

Heracleum crenatifolium Boiss. species accepted

Heracleum coreanum H.Boissieu species doubtful

Heracleum colchicum Lipsky species synonym

Heracleum circassicum Manden. species synonym

Heracleum chorodanum (Hoffm.) DC. species accepted

Heracleum caucasicum Steven species synonym

Heracleum caspicum DC. species synonym

Heracleum carmeli Wender. species synonym

Heracleum ×carbonnieri Reduron, Michaud & J.Molina species accepted

Heracleum calcareum Albov species synonym

Heracleum brevivittatum Ledeb. species synonym

Heracleum biternatum W.W.Sm. species accepted

Heracleum bipinnatum Guss. ex DC. species synonym

Heracleum bhutanicum M.F.Watson species accepted

Heracleum benearnense Rouy & E.G.Camus species synonym

Heracleum basvicum Fisch. species accepted

Heracleum barbatum Ledeb. species synonym

Heracleum bailletianum Timb.-Lagr. & Marcais species doubtful

Heracleum auritum Bisch. species synonym

Heracleum argaeum Boiss. & Balansa species accepted

Heracleum arcticum Rupr. species doubtful

Heracleum apiifolium Boiss. species accepted

Heracleum antasiaticum Manden. species accepted

Heracleum anisactis Boiss. & Hohen. species accepted

Heracleum anatolicum Manden. species synonym

Heracleum amplissimum Wender. species synonym

Heracleum amanum Boiss. & Kotschy species accepted

Heracleum alpinum Siev. species heterotypic synonym

Heracleum albovii Manden. species accepted

Heracleum akasimontanum Koidz. species accepted

Heracleum afghanicum Kitam. species synonym

Heracleum aestivum Jord. species synonym

Heracleum aconitifolium Woronow species accepted

Heracleum abyssinicum (Boiss.) C.Norman species accepted

Pastinaca aspera (Hoffm.) Koso-Pol. species synonym

Symphyoloma graveolens C.A.Mey. species synonym

Sphondylium villosum Hoffm. species synonym

Sphondylium subvillosum Hoffm. species synonym

Sphondylium speciosum Hoffm. species synonym

Sphondylium pubescens Hoffm. species synonym

Sphondylium marginatum Hoffm. species synonym

Sphondylium lanatum (Michx.) Nieuwl. & Lunell species synonym

Sphondylium gummiferum (Willd.) Hoffm. species synonym

Sphondylium giganteum Fisch. ex Hoffm. species synonym

Sphondylium asperum Hoffm. species synonym

Sphondylium alpinum (L.) Scop. ex Steud. species synonym

Wendia schelkovnikovii (Woronow) Grossh. species homotypic synonym

Wendia pastinacifolia (K.Koch) Grossh. species synonym

Wendia minima (Lam.) Bonnier & Layens species synonym

Wendia incana (Boiss. & A.Huet) Grossh. species synonym

Wendia hymenocoleon Woronow ex Grossh. species synonym

Wendia chorodanum Hoffm. species synonym

Wendia brevivittata (Ledeb.) Grossh. species synonym

Wendia alpina (L.) Schur species synonym

Peucedanum vocontiorum Spreng. species synonym

Peucedanum stolzii (Engl. & H.Wolff) M.Hiroe species synonym

Peucedanum rivae (Engl.) M.Hiroe species synonym

Heracleum zozimioides Benth. & Hook.f. species doubtful

Heracleum yungningense Hand.-Mazz. species accepted

Heracleum wolongense F.T.Pu & X.J.He species accepted

Heracleum wilhelmsii Fisch. & Avé-Lall. species accepted

Heracleum wenchuanense F.T.Pu & X.J.He species accepted

Heracleum voroschilovii Gorovoj species synonym

Heracleum viscosum Fisch. ex Steud., 1821 species accepted

Heracleum villosum (Hoffm.) Fisch. ex Spreng. species accepted

Heracleum vicinum H.Boissieu species accepted

Heracleum vescens Willd. ex Steud., 1821 species accepted

Heracleum verrucosum Steven ex Steud., 1840 species accepted

Heracleum umbonatum Boiss. species synonym

Heracleum umbellatum Gueldenst. species doubtful

Heracleum turugisanense Honda species synonym

Heracleum tuberosum Molina ex Willd. species doubtful

Heracleum transcaucasicum Manden. species synonym

Heracleum trachyloma Fisch. & C.A.Mey. species accepted

Heracleum tiliifolium H.Wolff species accepted

Heracleum testiculatum Lapeyr. species synonym

Heracleum tenuevittatum Dalla Torre & Sarnth. species doubtful

Heracleum taylorii C.Norman species accepted

Heracleum tabuchii Akasawa species synonym

Heracleum sumatranum Buwalda ex Steenis species accepted

Heracleum subvillosum Steven species synonym

Heracleum subglabrum Kitam. species accepted

Heracleum stenopteroides Fedde species homotypic synonym

Heracleum sphondylium var. tsurugisanense (Honda) H.Ohba variety synonym

Heracleum sphondylium f. rubriflorum H.Ohba form proparte synonym

Heracleum sphondylium var. nipponicum (Kitag.) H.Ohba variety synonym

Heracleum sphondylium f. dissectum H.Ohba form proparte synonym

Heracleum sphondylium subsp. cyclocarpum (K.Koch) P.H.Davis subspecies synonym

Heracleum sphondylium subsp. carpaticum (Porcius) Soó subspecies synonym

Heracleum stenopterum Diels species accepted

Heracleum sphondylium var. akasimontanum (Koidz.) H.Ohba variety synonym

Heracleum speciosum Hoffm. ex Link species synonym

Heracleum spathiphyllum K.Koch ex Boiss. species synonym

Heracleum souliei H.Boissieu species accepted

Heracleum sosnowskyi Manden. species accepted

Heracleum sommieri Manden. species accepted

Heracleum sisianense Boiss. & Buhse species synonym

Heracleum cyclocarpum var. ponticum Lipsky variety synonym

Heracleum carpaticum Porcius species accepted

Pastinaca dissecta (Ledeb.) Koso-Pol. species heterotypic synonym

Malabaila abyssinica var. kirungae (Engl.) Robyns variety synonym

Heracleum pumilum Vill. species accepted

Selinum alpinum (L.) E.H.L.Krause species synonym

Heracleum longifolium Steven species synonym

Heracleum barbatum subsp. moellendorffii (Hance) M.Hiroe subspecies synonym

Pastinaca incana (Boiss. & A.Huet) Koso-Pol. species heterotypic synonym

Heracleum schansianum Fedde ex H.Wolff species accepted

Heracleum panaces W.D.J.Koch, 1843 species accepted

Heracleum morifolium var. angustum (Nakai) Kitag. variety synonym

Heracleum speciosum Hornem. species synonym

Heracleum speciosum Weinm. species synonym

Heracleum pyrenaicum Bleb., 1808 species accepted

Heracleum acuminatum Franch. species heterotypic synonym

Heracleum speciosum Ledeb. species heterotypic synonym

Pastinaca schelkownikowii (Woronow) Koso-Pol. species synonym

Heracleum stevenii Manden. species synonym

Heracleum moellendorffii var. subbipinnatum (Franch.) Kitag. variety synonym

Heracleum pastinacifolium C.Koch species doubtful

Heracleum moellendorffii Hance species accepted

Heracleum leskovii Grossh. species accepted

Heracleum dulce Fisch., C.A.Mey. & Avé-Lall. species synonym

Heracleum dissectum Ledeb. species accepted

Heracleum moellendorffii f. angustum (Kitag.) Kitag. form synonym

Malabaila elgonense H.Wolff species synonym

Heracleum lescovii Grossh. species synonym

Heracleum persicum var. buhseanum Thell. variety synonym

Heracleum spathiphyllum K.Koch species homotypic synonym

Heracleum sphondylium f. subsetulosum Thell. form synonym

Heracleum alpinum subsp. juranum (Briq.) Thell. subspecies synonym

Heracleum mantegazzianum var. immaculatum Wilczek & Thell. variety synonym

Heracleum persicum x H. sibiricum species accepted

Heracleum pubescens var. persicum (Desf. ex Fisch., C.A.Mey. & Avé-Lall.) Regel variety synonym

Heracleum pubescens var. glabrum Regel variety synonym

Heracleum alpinum L. species accepted

Heracleum acutifolium Arv.-Touv., 1872 species synonym

Heracleum pastinacifolium K.Koch species accepted

Angelica oncosepala Hand.-Mazz. species synonym

Barysoma villosum (Hoffm.) Bunge species synonym

Malabaila stolzii Engl. & H.Wolff species synonym

Malabaila rivae Engl. species synonym

Malabaila kirungae Engl. species synonym

Malabaila elgonensis H.Wolff species synonym

Malabaila abyssinica Boiss. species synonym

Selinum wilhelmsii (Fisch. & Avé-Lall.) E.H.L.Krause species synonym

Pastinaca villosa (Hoffm.) Calest. species synonym

Pastinaca umbonata (Boiss.) Koso-Pol. species synonym

Pastinaca pubescens (Hoffm.) Calest. species synonym

Pastinaca pastinacella Koso-Pol. species synonym

Pastinaca pachyrhiza (Sommier & Levier) Koso-Pol. species synonym

Pastinaca orphanidis (Boiss.) Calest. species synonym

Pastinaca minima (Lam.) Calest. species synonym

Pastinaca ligusticifolia (M.Bieb.) Calest. species heterotypic synonym

Pastinaca lebmaniana Koso-Pol. species synonym

Pastinaca lanata (Michx.) Koso-Pol. species synonym

Pastinaca humilis (Sm.) Calest. species synonym

Pastinaca freyniana (Sommier & Levier) Koso-Pol. species synonym

Pastinaca cyclocarpa (K.Koch) Koso-Pol. species synonym

Pastinaca colchica (Lipsky) Koso-Pol. species synonym

Pastinaca chorodanum (Hoffm.) Koso-Pol. species synonym

Pastinaca caspica (DC.) Calest. species synonym

Pastinaca caroli-kochii Koso-Pol. species synonym

Pastinaca calcarea (Albov) Koso-Pol. species synonym

Pastinaca brevivittata (Ledeb.) Koso-Pol. species synonym

Pastinaca barbata (Ledeb.) Koso-Pol. species synonym

Pimpinella nandensis C.Norman species synonym

Pastinaca apiifolia (Boiss.) Koso-Pol. species synonym

Pastinaca alpina (L.) Calest. species synonym

Wendtia schelkownikowii Woronow species synonym

Wendtia argaea (Boiss. & Balansa) Woronow species synonym

Heracleum lanatum var. akasimontanum (Koidz.) H.Hara variety synonym

Heracleum incanum var. lazicum Boiss. variety synonym

Heracleum barmanicum Kurz species accepted

Heracleum pubescens var. laeve Freyn & Sint. variety synonym

Heracleum sphondylium subsp. alpinum (L.) Bonnier & Layens subspecies synonym

Heracleum apiifolium var. dissectum Freyn & Sint. variety synonym

Heracleum pubescens var. trachyloma (Fisch. & C.A.Mey.) Boiss. variety synonym

Heracleum microcarpum var. subbipinnatum Franch. variety synonym

Heracleum morifolium f. angustum Kitag. form synonym

Heracleum moellendorffii f. subbipinnatum (Franch.) Kitag. form synonym

Heracleum lanatum var. nipponicum (Kitag.) H.Hara variety synonym

Heracleum barbatum var. tsurugisanense (Honda) M.Hiroe variety synonym

Heracleum lanatum subsp. asiaticum M.Hiroe subspecies synonym

Heracleum sphondylium subsp. lanatum (Michx.) Á.Löve & D.Löve subspecies synonym

Heracleum lanatum var. asiaticum (M.Hiroe) H.Hara variety synonym

Heracleum tsurugisanense Honda species synonym

Heracleum moellendorffii var. tsurugisanense (Honda) Ohwi variety synonym

Heracleum mantegazzianum Sommier & Levier species accepted

Heracleum maximum W.Bartram species accepted

Heracleum lanatum Michx. species synonym

Heracleum sphondylium var. lanatum (Michx.) Dorn variety synonym

Heracleum sphondylium L. species accepted

Heracleum lecokii var. aurasiacum Maire variety synonym

Heracleum austriacum L. species accepted

Heracleum alpinum Georgi species synonym

Heracleum dissectum var. barbatum (Ledeb.) Krylov variety synonym

Heracleum montanum Schleich., 1821 species accepted

Heracleum adae Kulieva species doubtful

Heracleum lanatum subsp. moellendorffii (Hance) H.Hara subspecies synonym

Sphodylium pubescens Hoffm., 1814 species synonym

Heracleum sphondylium Bourg. ex Reut., 1864 species doubtful

Heracleum lanatum var. asiaticum Fisch. variety synonym

Heracleum dedenense R.H.Shan & Z.H.Pan species accepted

Sphondylium lanatum (Michx.) Greene species synonym

Heracleum sphondylium subsp. juranum (Genty) Thell. subspecies synonym

Heracleum armoricum Boreau ex Nyman species synonym

Heracleum giganteum Fisch. species synonym

Heracleum verticillatum Pančić ex Boiss., 1888 species accepted

Heracleum strelkovii Grossh. species synonym

Heracleum sphondylium var. tsaurugisanense (Honda) H.Ohba variety synonym

Sphondylium alpinum (L.) Caruel species synonym

Heracleum asperum (Hoffm.) M.Bieb. species accepted

Heracleum lanatum subsp. voroschilovii (Gorovoj) Stapf & Vorosch. subspecies synonym

Heracleum palmatum Arv.-Touv., 1871 species synonym

Pastinaca lehmanniana (Bunge) Koso-Pol. species synonym

Heracleum sphondylium Claus species doubtful

Heracleum panaces Steven species heterotypic synonym

Heracleum speciosum Rudolph species synonym

Heracleum viscosum Fisch. ex Spreng. species doubtful

Heracleum cyclocarpum K.Koch species accepted

Heracleum longifolium M.Bieb. species heterotypic synonym

Heracleum schelkovnikovii Woronow species accepted

Heracleum tromsoensis Elven species synonym

Heracleum pubescens var. wilhelmsii (Fisch. & Avé-Lall.) Boiss. variety synonym

Heracleum alpinum subsp. juranum (Genty) Rouy & E.G.Camus subspecies synonym

Heracleum sphondylium var. caucasicum Sommier & Levier variety synonym

Heracleum leucocarpum Rech.fil. species synonym

Heracleum mantegazzianum x sphondylium subsp. sphondylium species accepted

Heracleum persicum × sphondylium subsp. sibiricum species accepted

Heracleum mantegazzianum × sphondylium ssp. sphondylium species synonym

Heracleum panaces Pall. ex Hablitz species heterotypic synonym

Heracleum intermedium Ledeb. species heterotypic synonym

Heracleum stevenii Manden. ex Grossh., 1949 species accepted

Heracleum pastinacifolium subsp. schelkovnikovii (Woronow) Satzyp. subspecies homotypic synonym

Heracleum persicum Desf. ex Fisch., C.A.Mey. & Avé-Lall. species synonym

Heracleum woroschilowii Gorovoj species doubtful

Heracleum peshmenianum Ekim species accepted

Pastinaca mantegazziana (Sommier & Levier) Koso-Pol. species synonym

Heracleum stenopteroides Fedde ex H.Wolff species accepted

Heracleum pyrenaicum M.Bieb. species synonym

Heracleum angustifolium M.Bieb. species heterotypic synonym

Heracleum sphondylium Pall., 1773 species doubtful

Heracleum sphondylium Bourg. species homotypic synonym

Heracleum bipinnatum Guss. species homotypic synonym

Heracleum ponticum (Lipsky) Schischk. species homotypic synonym

Heracleum pulchellum Pall. species homotypic synonym

Heracleum marginatum Fisch. species homotypic synonym

Heracleum viscosum Fisch. species homotypic synonym

Heracleum vescens Willd. species homotypic synonym

Heracleum decipiens Fisch. species homotypic synonym

Heracleum grandiflorum Steven species homotypic synonym

Heracleum verrucosum Steven species homotypic synonym

Heracleum panaces Pall. species homotypic synonym

Heracleum pusillum Pall. species homotypic synonym

Heracleum montanum Georgi species homotypic synonym

Heracleum speciosum Hoffm. species homotypic synonym

Heracleum sumatranum Buwalda species homotypic synonym

Heracleum moellendorffii var. akasimontanum (Koidz.) Okuyama variety homotypic synonym

Sphondylium alpinum (L.) Scop. species homotypic synonym

Wendia hymenocoleon Woronow species homotypic synonym

Heracleum villosum (Hoffm.) Fisch. species homotypic synonym

Heracleum tuberosum Molina species homotypic synonym